Nutrient movement and mobility inside the plant:
Besides endocytosis, there are two major pathways inside the plant, the xylem and the phloem. When water and minerals are absorbed by plant roots, these substances must be transported up to the plant's stems and leaves for photosynthesis and further metabolic processes. This upward transport happens in the xylem. While the xylem is able to transport organic compounds, the phloem is much more adapted to do so.
The organic compounds thus originating in the leaves have to be moved throughout the plant, upwards and downwards, to where they are needed. This transport happens in the phloem. Compounds that are moving through the phloem are mostly:
Sugars as sugary saps, organic nitrogen compounds (amino acids and amides, ureides and legumes), hormones and proteins.
Not all nutrient compounds are moveable within the plant.
1) N, P, K, Mg and S are considered mobile: they can move up and down the plant in both xylem and phloem.
Deficiency appears on old leaves first.
2) Ca, Fe, Zn, Mo, B, Cu, Mn are considered immobile: they only move up the plant in the xylem.
Deficiency appears on new leaves first.
Storage organelles:
Salts and organic metabolites can be stored in storage organelles. The most important storage organelle is the vacuole, which can contribute up to 90% of the cell volume. The majority of compounds found in the vacuole are sugars, polysaccharides, organic acids and proteins though.
Translocation:
Now that the basics are explained, we can take a look at the translocation process. It should be already clear that only mobile elements can be translocated through the phloem. Immobile elements cant be translocated and are not more available to the plant for further metabolic processes and new plant growth.
Since flushing (in theory) induces a nutrient deficiency in the rootzone, the translocation process aids in the plants survival. Translocation is transportation of assimilates through the phloem from source (a net exporter of assimilate) to sink (a net importer of assimilate). Sources are mostly mature fan leaves and sinks are mostly apical meristems, lateral meristem, fruit, seed and developing leaves etc.
You can see this by the yellowing and later dying of the mature fan leaves from the second day on after flushing started. Developing leaves, bud leaves and calyxes don’t serve as sources, they are sinks. Changes in those plant parts are due to the deficient immobile elements which start to indicate on new growth first.
Unfortunately, several metabolic processes are unable to take place anymore since other elements needed are no longer available (the immobile ones). This includes processes where nitrogen and phosphorus, which have likely the most impact on taste, are involved.
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